grounds, the species range of those areas (Figure 6) is usually lower, nevertheless, than for both freshwater and totally marine environments, particularly in temperate waters. Those areas of estuaries, lagoons and coastal marshes, between purely freshwater and purely marine environments, signify an necessary but physiologically troublesome environment for many aquatic organisms. The concept of the trophic pyramid and the meals chain were developed in terrestrial ecology, where food linkages are typically more particular and less complicated than
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